Publications of the Group


Group publications are from 2016 on.


  • Gokhale, C.S., and Sharma, N. Leveraging Parrondo’s paradox for sustainable agriculture. ResearchSquare-rs-1141107/v1
  • Bargués-Ribera, M., Reeves, R.G., Gokhale, C.S., Eco-evolutionary dynamics of Plasmodium genotypes under mass drug administration, bioRxiv-818039
  • Rabajante, J.F., Anzia, E.L., Gokhale, C.S., On a mechanistic process of macroparasite aggregation, bioRxiv-680041
  • Gokhale, C.S., Wignall, A.E., On the innovation and evolution of predatory tactics, bioRxiv-530238


Journal articles



  1. Verma, P., Reeves, R. G., Simon, S. Otto, M. and Gokhale, C. S. 2021 The effect of mating complexity on gene drive dynamics. The American Naturalist forthcoming bioRxiv-2021.09.16.460618
  2. Gokhale, C. S., Bulbulia, J. & Frean, M. 2022 Collective narratives catalyse cooperation. Humanit Soc Sci Commun 9, 85 socRxiv
  3. Gao, Y., Pichugin, Y., Gokhale, C. S. & Traulsen, A. 2022 Evolution of reproductive strategies in incipient multicellularity. J Roy Soc Interface 19, 20210716. bioRxiv-2021.09.13.460035
  4. Traulsen, A., Gokhale, C.S., Shah, S. & Uecker, H. The Covid-19 pandemic: Basic insights from basic mathematical models, forthcoming NAL Live - Nationale Akademie Leopoldina

  5. Gokhale, C.S., Giamo, S., & Remigi, P. 2021 Memory shapes microbial populations, PLoS Computational Biology, 17, e1009431 bioRxiv-370106
  6. Verma, P., Reeves, R. G. & Gokhale, C. S. 2021 A common gene drive language eases regulatory process and eco-evolutionary extensions. BMC Ecol Evol 21, 156. bioRxiv-970103
  7. Mafessoni, F., Lachmann, M. & Gokhale, C. S. 2021 On the fitness of informative cues in complex environments. J Theor Biol 527, 110819 bioRxiv-066571
  8. Venkateswaran, V. R., Roth, O. & Gokhale, C. S. 2021 Consequences of combining sex‐specific traits. Evolution doi:10.1111/evo.14204. bioRxiv-version-892810
  9. Park, H.J.$, Gokhale, C.S.$ , Bertels, F. ($ joint first authors), 2021. How sequence populations persist inside bacterial genomes. Genetics doi:10.1093/genetics/iyab027.bioRxiv-version-170514

  10. Pathak, S., Verma, P., Ram, S. K. & Sengupta, S. How strategy environment and wealth shape altruistic behaviour: cooperation rules affecting wealth distribution in dynamic networks. Proc Royal Soc B, 287, 20202250 bioRxiv version-077131
  11. Denton, J.A., Gokhale, C.S., (joint first authors) 2020. Synthetic symbiosis under environmental disturbances. mSystems, 5 (3) bioRxiv version-395426
  12. Bargués-Ribera, M., Gokhale, C.S., 2020. Eco-evolutionary agriculture: host-pathogen dynamics in crop rotations. PLOS Computational Biology, 16(1): e1007546 bioRxiv-402313

  13. Venkateswaran, V.R., Gokhale, C.S., 2019. Evolutionary dynamics of complex multiple games. Proceedings of the Royal Society B: Biological Sciences,286(1905), p.20190900 OpenAccess bioRxiv-302265
  14. Denton, J.A., Gokhale, C.S., 2019. Synthetic mutualism and the intervention dilemma. Life - Special Issue “Synthetic Biology: From Living Computers to Terraformation”, 9 (1), 15
  15. Park, H-J., Gokhale, C.S., 2019. Ecological feedback on diffusion dynamics. Journal of the Royal Society Open Science, 6 (2), 181273

  16. Czuppon, P, Gokhale, C.S., 2018. Disentangling eco-evolutionary effects on trait fixation. Theoretical Population Biology, 124, 93-107.
  17. Roemhild, R., Gokhale, C.S., Dirksen, P., Blake, C., Rosenstiel, P., Traulsen, A., Andersson, D., Schulenburg, H., 2018. Cellular hysteresis as a novel principle to maximize the efficacy of antibiotic therapy, PNAS, 115(39), 9767-9772.

  18. Daubech, B., Remigi, P., Doin de Moura, G., Marchetti, M., Pouzet, C., Auriac, M-C., Gokhale, C.S., Masson-Biovin C., Capela, D., 2017. Spatio-temporal control of mutualism in legumes helps spread symbiotic nitrogen fixation, eLife, 6, 57.
  19. Schenk, H., Traulsen, A., Gokhale, C.S., 2017. Chaotic provinces in the kingdom of the Red Queen. Journal of Theoretical Biology, 431, pp.1–10.
  20. Liu, Y., Gokhale C.S., Rainey P.B., Zhang, X-X., 2017. Unravelling the complexity and redundancy of carbon catabolic repression in Pseudomonas fluorescens SBW25. Molecular microbiology, 105(4), pp.589–605.
  21. Bertels, F., Gokhale, C.S., Traulsen, A., 2017. Discovering Complete Quasispecies in Bacterial Genomes. Genetics, 206(4), pp.2149–2157.
  22. Gokhale, C.S., Traulsen, A., Joop, G., 2017. Social dilemma in the external immune system of the red flour beetle? It is a matter of time. Ecology & Evolution, 7(17), pp.6758–6765.

  23. Gokhale, C.S., Hauert, C., 2016. Eco-evolutionary dynamics of social dilemmas. Theoretical Population Biology, 111, 28–42.
  24. Papkou, A., Gokhale, C.S., Traulsen, A., Schulenburg, H., 2016. Host-parasite coevolution: why changing population size matters. Zoology (Jena, Germany), 119(4), pp.330–338.

    Publications of Gokhale prior to group formation
  25. Pichugin, Y., Gokhale, C.S., Garcia, J., Traulsen, A., Rainey, P.B., 2015. Modes of migration and multilevel selection in evolutionary multiplayer games. Journal of Theoretical Biology, 387, 144–153.
  26. Song, Y. Gokhale, C.S., Papkou, A., Schulenburg, H.,Traulsen, A., 2015. Host-parasite coevolution in populations of constant and variable size. BMC Evolutionary Biology, 15(1), p.212.
  27. Bauer, B., Gokhale, C.S., 2015. Repeatability of evolution on epistatic landscapes. Scientific reports, 5, p.9607.

  28. Gokhale, C.S., Traulsen, A., 2014. Evolutionary Multiplayer Games. Dynamic Games and Applications, 4(4), pp.468–488.
  29. Gokhale, C.S., Reeves, R.G., Reed, F.A., 2014. Dynamics of a combined medea-underdominant population transformation system. BMC Evolutionary Biology, 14(1), 98.

  30. Gokhale, C.S., Papkou, A., Traulsen, A., Schulenburg, H., 2013. Lotka-Volterra dynamics kills the Red Queen: population size fluctuations and associated stochasticity dramatically change host-parasite coevolution. BMC Evolutionary Biology, 13(1), p.254.
  31. Wu, B., Gokhale C.S., van Veelen, M., Wang, L., Traulsen, A., 2013. Interpretations arising from Wrightian and Malthusian fitness under strong frequency dependent selection. Ecology & Evolution, 3(5), pp.1276–1280.
  32. Wu, B., Traulsen, A., Gokhale, C.S., 2013. Dynamic Properties of Evolutionary Multi-player Games in Finite Populations. Games, 4(2), pp.182–199.

  33. Gokhale, C.S., Traulsen, A., 2012. Mutualism and evolutionary multiplayer games: revisiting the Red King. Proceedings of the Royal Society B: Biological Sciences, 279(1747), 4611–4616.
  34. Wu, B., Gokhale C.S., Wang, L., Traulsen, A., 2012. How small are small mutation rates? Journal of Mathematical Biology, 64(5), pp.803–827.
  35. Han, T.A., Traulsen, A., Gokhale, C.S., 2012. On equilibrium properties of evolutionary multi-player games with random payoff matrices. Theoretical Population Biology, 81(4), pp.264–272.

  36. Gokhale, C.S., Traulsen, A., 2011. Strategy abundance in evolutionary many-player games with multiple strategies. Journal of Theoretical Biology, 283, 180–191.

  37. Altrock, P.M., Gokhale, C.S., Traulsen, A., 2010. Stochastic slowdown in evolutionary processes. Physical Review E, 82(1), 11925.
  38. Gokhale, C.S., Traulsen, A., 2010. Evolutionary games in the multiverse. PNAS, 107(12), 5500–5504.

  39. Gokhale, C.S., Iwasa, Y., Nowak, M., Traulsen A., 2009. The pace of evolution across fitness valleys. Journal of Theoretical Biology, 259(3), 613–620.
  40. Bhardwaj, A., Mukerji, M., Sharma, S., Paul, J., Gokhale, C.S., Srivastava, A.K., Tiwari, S., 2009. MtSNPscore: a combined evidence approach for assessing cumulative impact of mitochondrial variations in disease., BMC Bioinformatics, 10 Suppl 8, S7.

Book Chapter


  1. Gokhale, C.S., Park, H-J., Eco-evolutionary Spatial dynamics of Non-linear Social Dilemmas, in Advances in Dynamic Games, eds. David M. Ramsey and Jérôme Renault bioRxiv version-660266

Book review

  • Gokhale, C.S. and Traulsen, A.T., Rethinking the origins of biological games, reviewing Game Theory in Biology: Concepts and Frontiers by John M. McNamara and Olof Leimar for the International Society for Behavioural Ecology newsletter