Publications of the Group


Group publications are from 2016 on.


  • Gokhale, C.S., Bulbulia, J., and Frean, M. Collective Narratives Catalyse Cooperation. SocArXiv. doi:10.31235/
  • Bargués-Ribera, M., Reeves, R.G., Gokhale, C.S., Eco-evolutionary dynamics of Plasmodium genotypes under mass drug administration, bioRxiv-818039
  • Rabajante, J.F., Anzia, E.L., Gokhale, C.S., On a mechanistic process of macroparasite aggregation, bioRxiv-680041
  • Gokhale, C.S., Wignall, A.E., On the innovation and evolution of predatory tactics, bioRxiv-530238


Book Chapter


  1. Gokhale, C.S., Park, H-J., Eco-evolutionary Spatial dynamics of Non-linear Social Dilemmas, in Advances in Dynamic Games, eds. David M. Ramsey and Jérôme Renault bioRxiv version-660266

Journal articles


  1. Gokhale, C.S., Giamo, S., & Remigi, P. 2021 Memory shapes microbial populations, PLoS Computational Biology, forthcoming bioRxiv-370106
  2. Verma, P., Reeves, R. G. & Gokhale, C. S. 2021 A common gene drive language eases regulatory process and eco-evolutionary extensions. BMC Ecol Evol 21, 156. bioRxiv-970103
  3. Mafessoni, F., Lachmann, M. & Gokhale, C. S. 2021 On the fitness of informative cues in complex environments. J Theor Biol 527, 110819 bioRxiv-066571
  4. Venkateswaran, V. R., Roth, O. & Gokhale, C. S. 2021 Consequences of combining sex‐specific traits. Evolution doi:10.1111/evo.14204. bioRxiv-version-892810
  5. Park, H.J.$, Gokhale, C.S.$ , Bertels, F. ($ joint first authors), 2021. How sequence populations persist inside bacterial genomes. Genetics doi:10.1093/genetics/iyab027.bioRxiv-version-170514

  6. Pathak, S., Verma, P., Ram, S. K. & Sengupta, S. How strategy environment and wealth shape altruistic behaviour: cooperation rules affecting wealth distribution in dynamic networks. Proc Royal Soc B, 287, 20202250 bioRxiv version-077131
  7. Denton, J.A., Gokhale, C.S., (joint first authors) 2020. Synthetic symbiosis under environmental disturbances. mSystems, 5 (3) bioRxiv version-395426
  8. Bargués-Ribera, M., Gokhale, C.S., 2020. Eco-evolutionary agriculture: host-pathogen dynamics in crop rotations. PLOS Computational Biology, 16(1): e1007546 bioRxiv-402313

  9. Venkateswaran, V.R., Gokhale, C.S., 2019. Evolutionary dynamics of complex multiple games. Proceedings of the Royal Society B: Biological Sciences,286(1905), p.20190900 OpenAccess bioRxiv-302265
  10. Denton, J.A., Gokhale, C.S., 2019. Synthetic mutualism and the intervention dilemma. Life - Special Issue “Synthetic Biology: From Living Computers to Terraformation”, 9 (1), 15
  11. Park, H-J., Gokhale, C.S., 2019. Ecological feedback on diffusion dynamics. Journal of the Royal Society Open Science, 6 (2), 181273

  12. Czuppon, P, Gokhale, C.S., 2018. Disentangling eco-evolutionary effects on trait fixation. Theoretical Population Biology, 124, 93-107.
  13. Roemhild, R., Gokhale, C.S., Dirksen, P., Blake, C., Rosenstiel, P., Traulsen, A., Andersson, D., Schulenburg, H., 2018. Cellular hysteresis as a novel principle to maximize the efficacy of antibiotic therapy, PNAS, 115(39), 9767-9772.

  14. Daubech, B., Remigi, P., Doin de Moura, G., Marchetti, M., Pouzet, C., Auriac, M-C., Gokhale, C.S., Masson-Biovin C., Capela, D., 2017. Spatio-temporal control of mutualism in legumes helps spread symbiotic nitrogen fixation, eLife, 6, 57.
  15. Schenk, H., Traulsen, A., Gokhale, C.S., 2017. Chaotic provinces in the kingdom of the Red Queen. Journal of Theoretical Biology, 431, pp.1–10.
  16. Liu, Y., Gokhale C.S., Rainey P.B., Zhang, X-X., 2017. Unravelling the complexity and redundancy of carbon catabolic repression in Pseudomonas fluorescens SBW25. Molecular microbiology, 105(4), pp.589–605.
  17. Bertels, F., Gokhale, C.S., Traulsen, A., 2017. Discovering Complete Quasispecies in Bacterial Genomes. Genetics, 206(4), pp.2149–2157.
  18. Gokhale, C.S., Traulsen, A., Joop, G., 2017. Social dilemma in the external immune system of the red flour beetle? It is a matter of time. Ecology & Evolution, 7(17), pp.6758–6765.

  19. Gokhale, C.S., Hauert, C., 2016. Eco-evolutionary dynamics of social dilemmas. Theoretical Population Biology, 111, 28–42.
  20. Papkou, A., Gokhale, C.S., Traulsen, A., Schulenburg, H., 2016. Host-parasite coevolution: why changing population size matters. Zoology (Jena, Germany), 119(4), pp.330–338.

    Publications of Gokhale prior to group formation
  21. Pichugin, Y., Gokhale, C.S., Garcia, J., Traulsen, A., Rainey, P.B., 2015. Modes of migration and multilevel selection in evolutionary multiplayer games. Journal of Theoretical Biology, 387, 144–153.
  22. Song, Y. Gokhale, C.S., Papkou, A., Schulenburg, H.,Traulsen, A., 2015. Host-parasite coevolution in populations of constant and variable size. BMC Evolutionary Biology, 15(1), p.212.
  23. Bauer, B., Gokhale, C.S., 2015. Repeatability of evolution on epistatic landscapes. Scientific reports, 5, p.9607.

  24. Gokhale, C.S., Traulsen, A., 2014. Evolutionary Multiplayer Games. Dynamic Games and Applications, 4(4), pp.468–488.
  25. Gokhale, C.S., Reeves, R.G., Reed, F.A., 2014. Dynamics of a combined medea-underdominant population transformation system. BMC Evolutionary Biology, 14(1), 98.

  26. Gokhale, C.S., Papkou, A., Traulsen, A., Schulenburg, H., 2013. Lotka-Volterra dynamics kills the Red Queen: population size fluctuations and associated stochasticity dramatically change host-parasite coevolution. BMC Evolutionary Biology, 13(1), p.254.
  27. Wu, B., Gokhale C.S., van Veelen, M., Wang, L., Traulsen, A., 2013. Interpretations arising from Wrightian and Malthusian fitness under strong frequency dependent selection. Ecology & Evolution, 3(5), pp.1276–1280.
  28. Wu, B., Traulsen, A., Gokhale, C.S., 2013. Dynamic Properties of Evolutionary Multi-player Games in Finite Populations. Games, 4(2), pp.182–199.

  29. Gokhale, C.S., Traulsen, A., 2012. Mutualism and evolutionary multiplayer games: revisiting the Red King. Proceedings of the Royal Society B: Biological Sciences, 279(1747), 4611–4616.
  30. Wu, B., Gokhale C.S., Wang, L., Traulsen, A., 2012. How small are small mutation rates? Journal of Mathematical Biology, 64(5), pp.803–827.
  31. Han, T.A., Traulsen, A., Gokhale, C.S., 2012. On equilibrium properties of evolutionary multi-player games with random payoff matrices. Theoretical Population Biology, 81(4), pp.264–272.

  32. Gokhale, C.S., Traulsen, A., 2011. Strategy abundance in evolutionary many-player games with multiple strategies. Journal of Theoretical Biology, 283, 180–191.

  33. Altrock, P.M., Gokhale, C.S., Traulsen, A., 2010. Stochastic slowdown in evolutionary processes. Physical Review E, 82(1), 11925.
  34. Gokhale, C.S., Traulsen, A., 2010. Evolutionary games in the multiverse. PNAS, 107(12), 5500–5504.

  35. Gokhale, C.S., Iwasa, Y., Nowak, M., Traulsen A., 2009. The pace of evolution across fitness valleys. Journal of Theoretical Biology, 259(3), 613–620.
  36. Bhardwaj, A., Mukerji, M., Sharma, S., Paul, J., Gokhale, C.S., Srivastava, A.K., Tiwari, S., 2009. MtSNPscore: a combined evidence approach for assessing cumulative impact of mitochondrial variations in disease., BMC Bioinformatics, 10 Suppl 8, S7.